I think it was running down the level of water ("mortgaging" the boiler) in order to keep the pressure up for a short period.
This was fine when climbing steeply uphill, as the water level would rise anyway at the back of the loco over the firebox (the important place) so it was straightforward to let it fall a bit. Mortgaging the boiler is actually using it as a power reservoir, consuming steam more rapidly than it is being generated.
This was however a nightmare on Dainton bank, west of Newton Abbot, where the top of the 1 in 36 up is immediately followed by 1 in 36 down to Totnes, something you don't get on the Lickey or elsewhere steep. Thank you Brunel! As a result you needed to come over the top with sufficient water level not to cause a problem as you tip over the top. The difference in water levels is 6" or more at the back of the boiler. I believe when the Britannias first came along, which had longer boilers than anything the GWR ever had, apart from The Great Bear (which was not allowed down to Plymouth) there was more than one case of a blown fusible plug at that point as a result.
Grierson the GWR chief civil engineer came up with a plan for deviation lines to overcome the issues of the South Devon banks, and it was put to the GW board for approval, all part of the many cut off main lines built 1900-10 on the GW. Churchward, who had a long-running disagreement with Grierson (Nock called it a feud), told the board he was able to design locomotives which could handle the banks for far less capital expenditure than the new lines and tunnels required. And so we are where we are there today.