455s are older than 321s and a similar age to 317s. Unless the financial climate changes drastically in the meantime I would expect to see the next long-term franchisees replacing them. The replacements may be suitable for use on Crossrail 2.
As to classes 317-322 and 455-456 not looking much like Mk3s, remember that the original ones had the same window depth as the Mk3 with some opening ventilators added just above as if they were an afterthought. The deeper windows with integral hoppers came in on later batches. Things like the internal panelling around the windows (cream GRP with black rubber spacers between them) are also very similar. I believe the basic Mk3 structure was used for all these units, obviously modified to be shorter, wider and with different door positions.