It is likely there will be some test / training runs over the full runs. When the Lowlanders were due to launch last October there were full "shadow" paths running behind the normal Lowlander just prior to the introduction that presumably were for "trial runs".
However, along the full routes this is likely to be the odd run or three, not weeks of continuous testing over every route. The main test routes were always from Polmadie to Arrochar & Tarbet on the WHL for north of Edinburgh / 73s (with several early test runs last year on this route) and then from Polmadie to Carlisle/Lancaster and various points in between for the WCML testing and mileage accumulation.
Just because Mk5s have not been to every point on the routes yet 3-4 months before the proposed launch doesn't mean it's becoming impossible for those dates to be achieved. Still plenty of time to fit in a small amount of physical test / trial runs to each of the destinations nearer the launch date. I suspect there's bigger challenges to deal with to achieve the latest launch dates, such as continuing to resolve any technical problems with the stock as they arise, training all the staff (train crew, hosts, guards, drivers, shunters, cleaners, maintenance teams etc etc); ensuring all the various shunting manoeuvres still work / can be accommodated etc etc.